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Psilotophyta

Psilotophyta

Psilotaceae
Tmesipteridaceae
Zosterophyllopsida† Psilotophyta (the "whisk ferns"; also sometimes as Psilophyta) is a division (i.e., phylum) of the Kingdom Plantae. This division contains only two genera, Psilotum, a small shrubby plant of the dry tropics, and Tmesipteris, an epiphyte found in Australia, New Zealand, and New Caledonia. There has long been controversy about the relationships of the Psilotophyta, with some claiming that they are ferns (Pteridophyta), and others maintaining that they are descendants of the first vascular plants. Recent evidence from DNA suggests a closer affinity to the ferns. All Psilotophyta share a few characteristics. Psilotophyta are vascular plants. They lack leaves, instead having small outgrowths called enations. The enations are not considered true leaves because there is only a vascular bundle just underneath them, but not inside, as in leaves. Psilotophyta also do not have true roots. They are anchored by rhizoids. Absorption is aided by symbiotic fungi called mycorrhizae. Three sporangia are united into a synangium, which is considered to be a very reduced series of branches. There is a thick tapetum to nourish the developing spores, as is typical of eusporangiate plants. The gametophyte looks like a small piece of subterranean stem, but produces antheridia and archegonia.

External links

- [http://www.ucmp.berkeley.edu/plants/pterophyta/psilotales.html Introduction to the Psilotales] sort21 Psilotophyta Category: cryptogams

Psilotum

- Psilotum nudum (L.) Beauvois
- Psilotum complanatum Sw. Psilotum (whisk ferns) is a genus of fern-like vascular plants, the sole genus in the family Psilotaceae and the order Psilotales. They have traditionally been thought not to be true ferns, but rather, odd "primitive" vascular plants that reproduce solely by spores, without seeds. Recent evidence has however suggested that they may in fact be ferns that have lost a number of pteridophytic characteristics, but their status is still uncertain. There are two species, Psilotum nudum and Psilotum complanatum, with a hybrid between them known, Psilotum x intermedium W. H. Wagner. The distribution of Psilotum is tropical and subtropical, in the New World, Asia, and the Pacific. The highest latitudes known are in South Carolina and southern Japan for P. nudum. Category:Plants ms:Paku-pakis whisk

Zosterophyllopsida

Zosterophyllopsida are an exinct family of ferns.

References

- [http://sn2000.taxonomy.nl/Main/Classification/199600.htm Classification from Systema Naturae 2000]

Division (biology)

:This article discusses categorisations of organisms. For a different meaning in biology, see cell division. In biology, the equivalent of a phylum in the plant or fungi kingdom is called a division. The main plant divisions, in the order in which they probably evolved, are the mosses (Division Bryophyta), the ferns (Division Filicophyta), the horsetails (Division Sphenophyta), the Cycads (Division Cycadophyta), the Ginkgo (Division Ginkgophyta), the conifers (Division Pinophyta), the Gnetophytes (Division Gnetophyta), and the angiosperms (Division Anthophyta). Angiosperms are the flowering plants that now dominate the plant world (80% of all vascular plants are angiosperms). rank02a th:ส่วน (ชีววิทยา)

Kingdom (biology)

In biology, a kingdom or regnum is the top-level, or nearly the top-level, taxon of organisms in scientific classification. Originally two kingdoms were distinguished: the Animalia for animals, and the Vegetabilia or Plantae for plants. Early authors also treated minerals in a third kingdom Mineralia. Each kingdom was divided into classes, later grouped into phyla for animals and divisions for plants. More recently, various other kingdoms have been created, and the ideal number and delineations are not settled. When single-celled organisms were first discovered, they were split between the two kingdoms: mobile forms in the animal phylum Protozoa, and colored algae and bacteria in the plant division Thallophyta or Protophyta. However, a number of forms were placed in both - for instance the mobile alga Euglena, and the amoeba-like slime moulds. As a result, Ernst Haeckel suggested creating a third kingdom Protista for them, although this was not very popular until relatively recently (sometimes as the Protoctista). The discovery that bacteria have a radically different cell structure from other organisms (prokaryotic rather than eukaryotic) led Herbert Copeland to give them a separate kingdom, originally called Mychota but later referred to as Monera or Bacteria. As it became apparent how important this distinction is, it became popular to divide living things into two superkingdoms or empires, called Prokaryota and Eukaryota. Whittaker recognized an additional kingdom for the Fungi. The resulting five-kingdom system, proposed in 1959, has become a popular standard and with some refinement is still used in many works, or forms the basis for newer multi-kingdom systems. It is based mainly on differences in nutrition: his Plantae were mostly multicellular autotrophs, his Animalia multicellular heterotrophs, and his Fungi multicellular saprotrophs. The remaining two kingdoms, Protista and Monera, included unicellular and simple cellular colonies. In the years around 1980 there was an emphasis on phylogeny and redefining the kingdoms to be monophyletic. The Animalia, Plantae, and Fungi were generally reduced to core groups of closely related forms, and the others thrown into the Protista. Based on rRNA studies Carl Woese divided the prokaryotes into two kingdoms, called Eubacteria and Archaebacteria. Such six-kingdom systems have become standard in many works. A variety of new eukaryotic kingdoms were also proposed, but most were quickly invalidated, ranked down to phyla or classes, or abandoned. The only one which is still in common use is the kingdom Chromista proposed by Cavalier-Smith, including organisms such as kelp, diatoms, and water moulds. Thus the eukaryotes are divided into three primarily heterotrophic groups, the Animalia, Fungi, and Protozoa, and two primarily photosynthetic groups, the Plantae (including red algae) and Chromista. However, it has not become widely used because of uncertainty over the monophyly of the latter two kingdoms. In 1990, Woese proposed that the Eubacteria, Archaebacteria, and Eukaryota represent three primary lines of descent and accordingly he promoted them to domains, renaming them Bacteria, Archaea, and Eucarya. This three-domain system has received notable criticism but has generally displaced the older two-empire system as a way of organizing kingdoms together. See also: Binomial nomenclature, Scientific classification, Taxonomy

References

- Whittaker, R.H. (1959). On the broad classification of organisms. Quart. Rev. Biol. 34: 210-226.
- Whittaker, R.H. (1969). New concepts of kingdoms of organisms. Science 163: 150-160.

Psilotum

Epiphyte

The term epiphyte refers to any plant that grows upon or attached to another living plant. The term stems from the Greek epi- (meaning 'upon') and phyton (meaning 'plant'). These plants are sometimes called "air plants" because they do not root in soil. However, there are many aquatic species of algae, including seaweeds, that are epiphytes on other aquatic plants (seaweeds or aquatic angiosperms). Of course there are also parasitic and semiparasitic plants growing on others. The best-known of this is the mistletoe. An epiphyte uses photosynthesis for energy and (where non-aquatic) obtains moisture from the air or from dampness (rain and cloud moisture) on the surface of its host. Roots may develop primarily for attachment, and specialized structures (for example, cups and scales) used to collect or hold moisture. Epiphytes are not parasitic on their host, growing independently and deriving only physical support. A clear advantage is gained over other herbs restricted to the ground (soil) surface where light is less available in the forest and herbivores are more active. However, epiphytes may grow so thickly as to damage the supporting plant. The best-known epiphytes include mosses, lichens, orchids, and bromeliads (such as Spanish moss and Tillandsia), but epiphytes may be found in every major group of the plant kingdom. Assemblages of large epiphytes occur most abundantly in moist tropical forests, but mosses and lichens occur as epiphytes in almost any environment with trees. In Europe one can find sometimes grass, small bushes and small trees growing in the forks of a branch. However there are no dedicated epiphytes using roots in Europe. The first important monograph on epiphyte ecology was written by A.F.W. Schimper (Die epiphytische Vegetation Amerikas, 1888). Epiphyte is one of the subdivisions of the Raunkiær system. ---- The fictitious company Epiphyte features prominently in Neal Stephenson's novel Cryptonomicon.

New Zealand

New Zealand or Aotearoa, the Land of the Long White Cloud, is a country of two large islands and many smaller islands in the south-western Pacific Ocean. New Zealand is notable for its isolation, being separated from Australia on the northwest by the Tasman Sea, some 2,000 km wide. The closest neighbours to the north are New Caledonia, Fiji and Tonga. The population of New Zealand is mostly of European descent, with the indigenous Māori as the largest minority. Non-Māori Polynesian and Asian peoples are also significant minorities, especially in the cities. Officially, Elizabeth II is the Queen of New Zealand and is represented in the country by a non-political Governor-General; though the Queen has no real political influence. Political power is held by the Prime Minister who is leader of the Government in the democratically elected Parliament of New Zealand. The monarch's Realm of New Zealand also includes the Cook Islands and Niue, which are entirely self-governing; Tokelau, which is moving towards self-government, and New Zealand's claim in Antarctica.

History

New Zealand is one of the most recently settled major land masses. Polynesian settlers arrived in their waka some time between 800 and 600 years ago to establish the indigenous Māori culture. Settlement of the Chatham Islands to the south-east of New Zealand produced the Moriori people but it is disputed whether they moved there from New Zealand or elsewhere in Polynesia. Most of New Zealand was divided into tribal territories called rohe, resources within which were controlled by an iwi ('tribe'). Usually no two iwi had overlapping rohe. Māori adapted to eating the local marine resources, flora and fauna for food, hunting the giant flightless moa (which soon became extinct), and ate the Polynesian Rat and kumara (sweet potato), which they introduced to the country. The first Europeans known to reach New Zealand were led by Abel Janszoon Tasman, who sailed up the west coast of the South and North islands in 1642. He named it Staten Landt, believing it to be part of the land Jacob Le Maire had discovered in 1616 off the coast of Chile. Staten Landt appeared on Tasman's first maps of New Zealand, but this was changed by Dutch cartographers to Nova Zeelandia, after the Dutch province of Zeeland, some time after Hendrik Brouwer proved the South American land to be an island in 1643. The Latin Nova Zeelandia became Nieuw Zeeland in Dutch. Lieutenant James Cook subsequently called the archipelago New Zealand, although the names he chose for the North and South islands were rejected, and the main three islands became known as North, Middle and South, with the Middle Island being later called the South Island. Cook began extensive surveys of the islands in 1769, leading to European whaling expeditions and eventually significant European colonisation. From as early as the 1780s, Māori had encounters with European sealers and whalers. Acquisition of muskets by those iwi in close contact with European visitors destabilised the existing balance of power between Māori tribes and there was a temporary but intense period of bloody inter-tribal warfare, known as the Musket Wars, that only ceased when all iwi were so armed. Concern about the exploitation of Māori by Europeans, Church Missionary Society lobbying and French interest in the region led the British to annex New Zealand by Royal Proclamation in January 1840. To legitimise the British annexation, Lieutenant Governor William Hobson had been dispatched in 1839; he hurriedly negotiated the Treaty of Waitangi with northern iwi on his arrival. The Treaty was signed in February, and in recent years it has come to be seen as the founding document of New Zealand. The Māori translation of the treaty promised the Māori tribes "tino rangatiratanga" would be preserved in return for cedeing kawanatanga, which the English versions translates as "chieftainship" for "sovereignty"; the real meanings are now disputed. Disputes over land sales and sovereignty caused the New Zealand land wars which took place between 1845 and 1872. In 1975 the Treaty of Waitangi Act established the Waitangi Tribunal, charged with hearing claims of Crown violations of the Treaty of Waitangi dating back to 1840. Some Māori tribes and the Moriori never signed the treaty. Although New Zealand was initially administered as a part of the Australian colony of New South Wales, it became a colony in its own right in 1841. European settlement progressed more rapidly than anyone anticipated, and settlers soon outnumbered Māori. Self-government was granted to the settler population in 1852. The first capital of New Zealand was Kororareka (known today as Russell) but shortly afterwards moved to Auckland. There were political concerns following the discovery of gold in Central Otago in 1861 that the South Island would form a separate colony. So in 1865 the capital was offically moved to the more central city of Wellington. New Zealand was involved in a Constitutional Convention in March 1891 in Sydney, New South Wales, along with the then-colonies of Australia. This was to consider a potential constitution for the proposed federation between the then-British Colonies of Australasia. New Zealand lost interest in joining Australia in a federation following this convention. New Zealand became an independent dominion on 26 September 1907 by royal proclamation. Full independence was granted by the United Kingdom Parliament with the Statute of Westminster in 1931; it was taken up upon the Statute's adoption by the New Zealand Parliament in 1947. Since then New Zealand has been a sovereign constitutional monarchy within the Commonwealth of Nations.

Politics

New Zealand is a constitutional monarchy with a parliamentary democracy. Under the New Zealand Royal Titles Act (1953), Queen Elizabeth II is Queen of New Zealand and is represented as head of state by the Governor-General, Dame Silvia Cartwright. The New Zealand Parliament has only one chamber, the House of Representatives which usually seats 120 members of Parliament. Parliamentary elections are every three years under a form of proportional representation called Mixed Member Proportional (MMP). The 2005 General Election created an 'overhang' of one extra seat (occupied by the Māori Party), due to that party winning more seats in constituencies than its proportional entitlement. There is no single written constitution; however, the Constitution Act (1986) is the principal formal statement of New Zealand's constitutional structure. The Governor-General has the power to appoint and dismiss Prime Ministers and to dissolve Parliament. The Governor-General also chairs the Executive Council which is a formal committee consisting of all ministers of the Crown. Members of the Executive Council are required to be members of Parliament, and most are also in Cabinet. Cabinet is the most senior policy-making body and is led by the Prime Minister who is also the Parliamentary leader of the governing party or coalition. The current Prime Minister is Helen Clark of the Labour Party. She has served two complete terms as Prime Minister and has begun her third. On 17 October 2005 she announced that she had come to a complex arrangement that guaranteed the support of enough parties for her Labour-led coalition to govern. The core of the coalition is a cabinet consisting of Labour Party ministers and Jim Anderton, the Progressive Party's only MP. In addition to the parties represented in cabinet the leaders of New Zealand First and United Future are to be appointed as Ministers outside Cabinet. An arrangement of this kind has never been attempted before in New Zealand. A further arrangement has been made with the Green Party, which has given a commitment not to vote against the government on confidence and supply. This commitment assures the government of a majority of seven MPs on confidence. The Leader of the Opposition is National Party leader Don Brash who was formerly Governor of the Reserve Bank. Also in opposition are the Māori Party and ACT New Zealand. The highest court in New Zealand is the Supreme Court of New Zealand. The Supreme Court was established in 2004 following the passage of the Supreme Court Act in 2003. The Act abolished the option to appeal Court of Appeal rulings to the Privy Council in London. The current Chief Justice is Dame Sian Elias. New Zealand's judicary also has a High Court which deals with serious criminal offences and civil matters, and a Court of Appeal, as well as subordinate courts.

Foreign relations and military

New Zealand maintains a strong profile on environmental protection, human rights and free trade, particularly for agriculture. New Zealand is a member of the following geo-political organisations: APEC, Commonwealth of Nations, OECD and the United Nations. It has signed up to a number of free trade agreements, of which the most important is Closer Economic Relations with Australia. For its first hundred years, New Zealand followed Britain's lead on foreign policy. "Where she goes, we go, where she stands, we stand", said Prime Minister Michael Savage, in declaring war on Germany on 3 September 1939. However, Britain's inability to protect New Zealand from Japanese aggression in World War II led New Zealand to come under the influence of the United States of America for the generation following the war. New Zealand has traditionally also worked closely with Australia, whose foreign policy followed a similar historical trend. In turn, many Pacific Islands such as Western Samoa have looked to New Zealand's lead. The American influence on New Zealand was weakened by the disappointment with the Vietnam War, the nuclear danger presented by the Cold War, the Sinking of the Rainbow Warrior by France and by disagreements over environmental and agricultural trade issues. New Zealand is a party to the ANZUS security treaty between Australia, New Zealand and the United States. In 1984 New Zealand refused nuclear-powered or nuclear-armed ships access to its ports. In 1986 the United States announced that it was suspending its treaty security obligations to New Zealand pending the restoration of port access. The New Zealand Nuclear Free Zone, Disarmament and Arms Control Act of 1987 prohibits the stationing of nuclear weapons on the territory of New Zealand and the entry into New Zealand waters of nuclear armed or propelled ships. This legislation remains a source of contention and the basis for the United States' continued suspension of treaty obligations to New Zealand. In addition to the various wars between Iwi, and between the British, settlers and Iwi, New Zealand has fought in the Boer War, World War I, (sustaining the highest casualties per head of population of any combatant nation), World War II, the Korean War, the Malayan Emergency (and committed troops, fighters and bombers to the subsequent confrontation with Indonesia), the Vietnam War, the Gulf War and the Afghanistan War and has briefly sent a unit of army engineers to help with rebuilding Iraqi infrastructure. The New Zealand military has three branches: the New Zealand Army, the Royal New Zealand Navy, and the Royal New Zealand Air Force. New Zealand considers its own national defence needs to be modest; it dismantled its air combat capability in 2001. New Zealand has contributed forces to recent regional and global peacekeeping missions, including those in Cyprus, Somalia, Bosnia, the Sinai, Angola, Cambodia, the Iran/Iraq border, Bougainville and East Timor.

Local government and external territories

East Timor The early European settlers divided New Zealand into provinces. These were abolished in 1876 so that government could be centralised for financial reasons. As a result, New Zealand has no separately represented subnational entities such as provinces, states or territories apart from its local government. The spirit of the provinces however still lives on, and there is fierce rivalry exhibited in sporting and cultural events. Since 1876, local government has administered the various regions of New Zealand. In 1989, the government completely reorganised local government, implementing the current two-tier structure of regional councils and territorial authorities. Today New Zealand has 12 regional councils for the administration of environmental and transport matters and 74 territorial authorities that administer roading, sewerage, building consents, and other local matters. The territorial authorities are 16 city councils, 57 district councils, and the Chatham Islands County Council. Four of the territorial councils (one city and three districts) and the Chatham Islands County Council also perform the functions of a regional council and thus are known as unitary authorities. Territorial authority districts are not subdivisions of regional council districts, and a few of them straddle regional council boundaries. Regions are (asterisks denote unitary authorities): Northland, Auckland, Waikato, Bay of Plenty, Gisborne
- , Hawke's Bay, Taranaki, Manawatu-Wanganui, Wellington, Marlborough
- , Nelson
- , Tasman
- , West Coast, Canterbury, Otago, Southland, Chatham Islands
- . As a major South Pacific nation, New Zealand has a close working relationship with many of the smaller Pacific Island nations, and continues a political association with the Cook Islands, Niue, and Tokelau. New Zealand operates Scott Base in its Antarctic territory, the Ross Dependency. Other countries also use Christchurch to support their Antarctic bases and the city is sometimes known as the "Gateway to Antarctica".

Geography

Ross Dependency are visible in the centre of the North Island. The Southern Alps and the rain shadow they create are clearly visible on the South Island]] New Zealand comprises two main islands (simply called the North and South Islands in English, or usually Te-Ika-a-Maui and Te Wai Pounamu in Māori) and a number of smaller islands. The total land area of New Zealand, 268,680 km², is a little less than that of Japan and a little more than the United Kingdom. The country extends more than 1600 km along its main, north-north-east axis. The most significant of the smaller inhabited islands of New Zealand include Stewart Island/Rakiura, Waiheke Island, an island in Auckland's Hauraki Gulf, Great Barrier Island, east of the Hauraki Gulf and the Chatham Islands, named Rekohu by Moriori. The country has extensive marine resources, with the fifth largest Exclusive Economic Zone in the world covering over 4 million km², more than 15 times its land area. The South Island is the largest land mass, and is divided along its length by the Southern Alps, the highest peak of which is Aoraki/Mount Cook, at 3,754 metres (12,316 feet). There are 18 peaks of more than 3,000 metres in the South Island. The North Island is less mountainous than the South, but is marked by volcanism. The tallest North Island mountain, Mount Ruapehu (2,797 metres), is an active cone volcano. The dramatic and varied landscape of New Zealand has made it a popular location for the production of television programmes and films, including the Lord of the Rings trilogy. Lord of the Rings The usual climate throughout the country is mild, mostly cool temperate to warm temperate, with temperatures rarely falling below 0°C or rising above 30°C. Conditions vary from wet and cold on the West Coast of the South Island to dry and continental in the Mackenzie Basin of inland Canterbury and subtropical in Northland. Of the main cities, Christchurch is the driest, receiving only some 640 millimetres of rain per year. Auckland, the wettest, receives a little less than three times that amount.

Flora and fauna

Northland Because of its long isolation from the rest of the world, and its island biogeography New Zealand has extraordinary flora and fauna. About 80 percent of the New Zealand flora only occurs in New Zealand, including more than 40 endemic genera. The main two types of forest have been dominated by podocarps including the giant kauri and southern beech. The remaining vegetation types in New Zealand are grassland of grass and tussock, usually associated with the subalpine areas, and the low shrublands between grasslands and forests. Until the arrival of the first humans, 80% of the land was forested and, barring two species of bat, there were no non-marine mammals at all. Instead, New Zealand's forests were inhabited by a diverse range of birds including the flightless Moa which is now extinct, the Kiwi, Kakapo, and Takahē which are all endangered due to human actions. Unique birds capable of flight include the Haast's eagle which was the world's largest bird of prey before it became extinct and the large parrots the Kaka and Kea. Reptiles present in New Zealand include skinks and geckos and the Tuatara. There are no snakes but there are many species of insects— including the weta which may grow as large as a House Mouse.

Economy

House Mouse New Zealand has a thriving, modern, developed economy. The country has a high standard of living, ranking 19th on the 2005 Human Development Index and 15th of The Economists 2005 world-wide quality-of-life index. Since 1984 successive governments have engaged in major macroeconomic restructuring, transforming New Zealand from a highly protectionist and regulated economy to a liberalised free-trade economy. During the late 1980s, the New Zealand Government sold a number of major trading enterprises, including its telecommunications company, railway network, a number of radio stations and two financial institutions in a series of asset sales. Although the New Zealand Government continues to own a number of significant businesses, collectively known as State-Owned Enterprises (SOEs), they are operated through arms-length shareholding arrangements as stand-alone businesses that are required to operate profitably, just like any privately owned enterprise. Unfortunately, due in part to the sudden transition to a market economy, an economic bubble developed in the New Zealand stock market starting in 1984. This burst in October 1987 and the total value of the market halved within a year (it has still to recover this lost value). The effect of this bubble was a period of poor economic growth which lasted until the mid 90s. It also led the government to begin a programme of massive immigration to boost GDP. However, since 1999 New Zealand has enjoyed a period of relatively strong and sustained growth, and contained inflationary pressures. The current New Zealand government's economic objectives are centred around moving from being ranked among the lower end of the OECD countries to regaining a higher placing again, pursuing free-trade agreements, "closing the gaps" between ethnic groups, and building a "knowledge economy." In 2004 it began discussing free trade with China, one of the first countries to do so. New Zealand is heavily dependent on trade—particularly in agricultural products—to drive growth, and it has been affected by global economic slowdowns and slumps in commodity prices. Since agricultural exports are highly sensitive to currency values and a large percentage of consumer goods are imported, any changes in the value of the New Zealand dollar has a strong impact on the economy. Its primary export industries are agriculture, horticulture, fishing, forestry and information technology. There are also substantial tourism and export education industries. The film and wine industries are considered to be up-and-coming.
Demographics
New Zealand has a population of about 4.1 million. About 70% of the population are whites of European descent. New Zealanders of Anglo-Celtic ancestry are known as Pākeha - this term is used variously and some Māori use it to refer to all non-Māori New Zealanders. A large proportion of white New Zealanders are of Scottish ancestry. Māori people are the second largest ethnic group (the percentage of the population of full or part-Māori ancestry is 14.7%; those who checked only Māori are 7.9%). Between the 1996 and 2001 censuses, the number of people of Asian origin (6.6%) overtook the number of people of Pacific Island origin (6.5%) (note that the census allowed multiple ethnic affiliations). Virtually all Māori are of mixed heritage (Māori/Pākeha), but a large portion of them marked themselves as Māori-only on the Census. New Zealand is positive about immigration and is committed to increasing its population by approx 1% per annum. At present migrants from the UK constitute the largest single group (30%) but new migrants are drawn from many nations, increasingly from East Asia. Christianity is the predominant religion in New Zealand, although nearly 40% of the population has no religious affiliation. The main Christian denominations are Anglicanism, Presbyterianism, Roman Catholicism and Methodism. There are also significant numbers who identify themselves with Pentecostal and Baptist churches and with the Mormon church. The New Zealand-based Ratana church has many adherents among Māori. According to census figures, other significant minority religions include Hinduism, Buddhism and Islam.
Culture
IslamNew Zealand has a diverse contemporary culture with influences from British, the Māori,and other European immigrants and most recently Polynesian cultures. There were many people from Scotland amongst the early British settlers and elements of their culture persist; New Zealand is said to have more bagpipe bands than Scotland. Cultural links between New Zealand and the UK are maintained by a common language, sustained migration from the UK and the fact that many young New Zealanders spend time in the UK on their "overseas experience (OE)". Pre-European contact Māori culture had no metal tools, relying on stone and wood. Modern Māori do not live a traditional lifestyle. Elements of Māori culture survive and the Government actively promotes it to all New Zealanders. Use of the Māori language (Te Reo Māori) as a living, community language remained only in a few remote areas in the post war years but it is currently going through a renaissance; with generous state support for Māori language medium schools and a Māori language television channel. New Zealand's landscape has appeared in a number of television programmes and films. In particular, the television series Hercules and Xena were filmed around Auckland, and the film Heavenly Creatures in Christchurch. The television series The Tribe is set and filmed in New Zealand as well. Director Peter Jackson shot the epic The Lord of the Rings trilogy in various locations around the country, taking advantage of the spectacular and relatively unspoiled landscapes, and Mount Taranaki was used as a stand-in for Mount Fuji in The Last Samurai. The latest of such major international films to be released are King Kong and The Chronicles of Narnia: The Lion, the Witch and the Wardrobe.
Sport
New Zealand's most popular sports are rugby union, cricket, netball, lawn bowling, soccer (the most popular sport amongst children) and rugby league. Also popular are golf, tennis, cycling and a variety of water sports, particularly sailing, whitewater kayaking, Surf Lifesaving and rowing. In the latter, New Zealand enjoyed an extraordinary Magic 45 minutes when winning four successive gold medals at the 2005 world championships. Snow sports such as skiing and snowboarding are also popular. Equestrian sportsmen and sportswomen make their mark in the world (Mark Todd being chosen international "Horseman of the Century"), and all the way down to the juniors at pony club level.
Olympic Games
The country is internationally recognised as achieving extremely well on a medals-to-population ratio at Olympic Games and Commonwealth Games. See, for example, New Zealand Olympic medallists and New Zealand at the 2004 Summer Olympics.
Rugby
Rugby as a sport is closely linked to New Zealand's national identity. The national rugby team is called the All Blacks and has the best winning record of any national team in the world, including being the inaugural winners of the World Cup in 1987. The style of name has been followed in naming the national team in several other sports. For instance, the nation's basketball team is known as the Tall Blacks. New Zealand is to host the 2011 Rugby Union World Cup. New Zealand's national sporting colours are not the colours of its flag, but are black and white (silver). The silver fern is a national emblem worn by New Zealanders representing their country in sport. The haka—a traditional Māori challenge—is often performed at sporting events. The All Blacks traditionally perform a haka before the start of international matches.
Yachting, America's Cup
New Zealand is one of the leading nations in world yachting, especially open water long distance or around the world races. Round-the-world yachtsman, Sir Peter Blake was something of a national hero before his untimely death at the hands of river pirates while on an environmental exploration trip on the Amazon. In inshore yachting, Auckland hosted the last two America's Cup regattas (2000 and 2003). In 2000, Team New Zealand successfully defended the trophy they had won in 1995 in San Diego, which made them the only team in the history of the Cup to successfully defend a challenge other than a United States team, but in 2003 they lost to a team headed by Ernesto Bertarelli of Switzerland, whose Alinghi syndicate was skippered by Russell Coutts, the former skipper of Team New Zealand. Team New Zealand will compete for the America's Cup at the next regatta in Valencia in 2007. The team manager is Grant Dalton.
Public holidays
Statutory Holidays
(These holidays are legislated by several Acts of Parliament, such as the Holidays Act. New Zealand Statutes can be viewed at [http://www.legislation.govt.nz/browse_vw.asp?content-set=pal_statutes legislation.govt.nz]) There are also Provincial Anniversary Days to celebrate the founding days or landing days of the first colonists of the various colonial provinces. The actual observance of Anniversary days can vary even within each province due to local custom, convenience or the proximity of seasonal events or other holidays. This may differ from the historical observance day, and may be several weeks from the historic date of the events being commemorated. A full list of Anniversary days is listed in the article Holidays in New Zealand.
International rankings

- UN Human Development Index (HDI), 2005: 19th out of 177 behind Norway; United Nations Development Programme (pdf) [http://hdr.undp.org/reports/global/2005/pdf/HDR05_complete.pdf]
- Quality of Life Index, 2005: 15th out of 111 behind Ireland; The Economist Intelligence Unit (pdf) [http://www.economist.com/media/pdf/QUALITY_OF_LIFE.pdf]
- Environmental Sustainability Index, 2005: 14th (out of 146) behind Finland; Yale University Center for Environmental Law and Policy & Columbia University Center for International Earth Science Information Network (pdf) [http://www.yale.edu/esi/ESI2005_Main_Report.pdf]
- Index of Economic Freedom, 2005: 5th= (out of 155) behind Hong Kong; Heritage Foundation/Wall Street Journal [http://www.heritage.org/research/features/index/]
- GDP Ranking, 2005: 25th out of 111 behind Luxembourg; The Economist Intelligence Unit (pdf) [http://www.economist.com/media/pdf/QUALITY_OF_LIFE.pdf]
- Transparency International 2005: 2nd= (out of 159) behind Iceland on its list of least corrupt countries in the world. [http://ww1.transparency.org/cpi/2005/cpi2005_infocus.html#cpi]
See also

Notes
McGlone, S.M. and Wilmshurst, J.M. 1999. Dating initial Māori environmental impact in New Zealand. Quaternary International 59:5 - 16
Ministry for the Environment. 2005. Offshore Options: Managing Environmental Effects in New Zealand's Exclusive Economic Zone. [http://www.mfe.govt.nz/publications/water/offshore-options-jun05/html/page3.html Introduction]
Allan, H.H. 1982. Indigenous Tracheophyta - Psilopsida, Lycopsida, Filicopsida, Gymnospermae, Dicotyledons, Flora of New Zealand Volume I. Botany Division, Department of Scientific and Industrial Research

External links

- [http://www.teara.govt.nz/ Te Ara, the Encyclopedia of New Zealand]
- [http://webdirectory.natlib.govt.nz/index.htm Te Puna Web Directory ] - A directory to New Zealand web sites
- [http://www.mch.govt.nz/ Ministry for Culture and Heritage] - includes information on flag, anthems and coat of arms.
- [http://www.govt.nz/ New Zealand Government Portal]
- [http://www.nzte.govt.nz/ New Zealand Trade and Enterprise] - New Zealand's trade and economic development agency.
- [http://wikitravel.org/en/article/New_Zealand Wikitravel] - New Zealand travel guide.
- [http://www.metservice.co.nz/ New Zealand weather]
- [http://www.nzhistory.net.nz/ NZHistory.net.nz New Zealand history website]
- [http://www.stats.govt.nz/ Statistics New Zealand] - Official statistics.
- [http://www.newzealand.com/ Tourism New Zealand]
- [http://www.astronomy.net.nz/ Astronomy in New Zealand] The guide to astronomy in New Zealand
- [http://www.nzherald.co.nz/ New Zealand Herald] - New Zealand newspaper online A
Category:Island nations Category:Members of the Commonwealth of Nations Category:Former British colonies Category:Monarchies Category:Oceanic countries Category:Polynesia als:Neuseeland zh-min-nan:Aotearoa ko:뉴질랜드 ms:New Zealand ja:ニュージーランド simple:New Zealand th:ประเทศนิวซีแลนด์

New Caledonia

:For the former North American colony called New Caledonia, see New Caledonia (Canada). New Caledonia (Canada) New Caledonia (French: Nouvelle-Calédonie; popular names: Kanaky, Le caillou) is a "sui generis collectivity" (in practice an overseas territory of France) made up of a main island (Grande-Terre de la Nouvelle-Calédonie) and several smaller islands. It is located in the region of Melanesia in the southwest Pacific. It has a land area of 18,575.5 km² (7,172 sq. miles). The population at the 2004 census was 230,789 inhabitants. It has an Internet country code top-level domain (ccTLD) of .nc. The capital and largest city of the territory is Nouméa. The currency is the Comptoirs Francais du Pacifique franc (international code XPF). Since 1986 the United Nations Committee on Decolonization has included New Caledonia on the United Nations list of Non-Self-Governing Territories, a highly political list that is disputed by France, Morocco, the United States, and the United Kingdom, which all have territories on the list. The list is partial and controversial because it does not cover all peoples and territorial entities in the world which are neither sovereign countries nor part of any sovereign country. New Caledonia will decide whether to remain within the French Republic or become an independent state in a referendum sometime after 2014. Its capital Nouméa is the seat of the regional organization the Secretariat of the Pacific Community (formerly the South Pacific Commission). Secretariat of the Pacific Community Secretariat of the Pacific Community]

Name

The name Caledonia derives from the Latin name of an area corresponding to modern Scotland. The name Kanaky is also in common usage in French, English and the indigenous languages. This name is favored by Melanesian nationalists. The word comes from kanaka, a Polynesian word meaning human used by Polynesians to call themselves. The word was later used by the French to call all the native inhabitants of the South Pacific Ocean, including the Melanesian (non-Polynesian) native inhabitants of New Caledonia. The word, turned into Canaque in French, became derogative. In the 1960s and 1970s, when the Melanesian native inhabitants started to organize themselves into political parties and ask for independence, this derogative word was turned into a symbol of political emancipation and pride. In 1983, during the period of political turmoil, the terms KANAK and KANAKY became a political brand names and almost instantly disappeared from the lips of the Caldoches who realised how the name had changed into a political statement.

Geography

Caldoche Main article: Geography of New Caledonia New Caledonia is located around in the southwest Pacific Ocean, approximately 1,200 km east of Australia and 1,500 km northwest of New Zealand. The island nation of Vanuatu lies to the northeast. New Caledonia is made up of a main island, the Grande Terre, and several smaller islands, the Belep archipelago to the north of the Grande Terre, the Loyalty Islands to the east of the Grande Terre, the Île des Pins to the south of the Grande Terre, and the Chesterfield Islands and Bellona Reefs further to the west. The Grande Terre is by far the largest of the islands, and the only mountainous island. It has an area of 16,372 km² (6,321 sq. miles), and is enlongated northwest-southeast, 350 km in length and 50 to 70 km wide. A mountain range runs the length of the island, with five peaks over 1500 meters. The highest point is Mont Panié at 1,628 meters elevation (5,341 ft). New Caledonia contains about one-quarter of the world's nickel resources.

Climate

New Caledonia lies astride the Tropic of Capricorn, between 19° and 23° south latitude. The climate of the islands is tropical, and rainfall is highly seasonal, brought by trade winds that usually come from the east. Rainfall averages about 1,500 mm yearly on the Loyalty Islands, 2,000 mm at low elevations on eastern the Grande Terre, and 2,000-4,000 mm at high elevations on the Grande Terre. The western side of the Grande Terre lies in the rain shadow of the central mountains, and rainfall averages 1200 mm per year.

Ecology

Unlike many of the Pacific islands which are of relatively recent volcanic origin, New Caledonia is an ancient fragment of the Gondwana supercontinent. New Caledonia and New Zealand separated from Australia 85 million years ago, and from one another 55 million years ago, and New Caledonia still carries many unique and endemic plants and animals of Gondwanan origin. (see Biodiversity of New Caledonia and Endemic Birds of New Caledonia). The best known is a hen-sized bird, the Cagou or Kagu, which cannot fly, has a large crest, and a funny cooing, and whose song and image serves as an emblem. The Niaouli tree, which also grows in Australia and New Guinea, is of medical interest, as its sap gives gomenol, which smells like camphor and is used to treat head colds. Before the Europeans arrived, there was no mammal other than the roussette (aka flying fox), a big vegetarian bat, a local delicacy. The islands make up two terrestrial ecoregions, the New Caledonia rain forests on the Loyalty Islands, Ile des Pins, and the eastern side of Grand Terre, and the New Caledonia dry forests in the rain shadow on the western side of Grand Terre. As the Europeans settled on the dry west coast and left the east to Kanaks, the political division maps the natural one. New Caledonia's freshwater ecology also evolved in long isolation, and the New Caledonia rivers and streams are home to many endemic species. The New Caledonia Barrier Reef, which surrounds the Grande Terre and the Île des Pins, is the second-largest coral reef in the world after Australia's Great Barrier Reef, reaching a length of 1500 km. The reef has great species diversity, is home to endangered dugongs (Dugong dugon), and is an important nesting site for Green Sea Turtle (Chelonia mydas).

Administration

Along with other Pacific Ocean's territories of French Polynesia and Wallis and Futuna, New Caledonia is part of the French Republic. Its official status is that of a sui generis collectivity (collectivité sui generis), a status unique in the French Republic. New Caledonia was a colony until 1946, then an overseas territory (territoire d'outre-mer, or TOM) from 1946 to 1999. The capital is Nouméa.

History

The western Pacific was first populated about 50,000 years ago. The Austronesians moved into the area later. The diverse group of people that settled over the Melanesian archipelagos are known as the Lapita. They arrived in the archipelago now commonly known as New Caledonia and the Loyalty Islands around 1500 BCE. The Lapita were highly skilled navigators and agriculturists with influence over a large area of the Pacific. From about the 11th century CE Polynesians also arrived and mixed with the populations of the archipelago. Europeans first sighted New Caledonia and the Loyalty Islands in the late 18th century. The British explorer James Cook sighted Grande Terre in 1774 and named it New Caledonia after the Scottish Highlands, which it resembled. Caledonia was a popular poetic and patriotic term for Scotland, and James Cook's father was Scottish. British and North American whalers and sandalwood traders became interested in New Caledonia and tensions developed as their approach became increasingly dishonest (an arrogant attitude and cheating became commonplace). Europeans used alcohol and tobacco amongst other things to barter for commodities. Contact with Europeans brought new diseases such as smallpox, measles, dysentery, influenza, syphilis and leprosy. Many people died as a result of these diseases. Tensions developed into hostilities and in 1849 the crew of the Cutter were killed and eaten by the Pouma clan. As trade in sandalwood declined it was replaced by a new form of trade. Blackbirding involved enslaving people from New Caledonia, the Loyalty Islands, Vanuatu, Papua New Guinea and the Solomon Islands to work in sugar cane plantations in Fiji and Queensland. The trade ceased at the start of the 20th century. Catholic and Protestant missionaries first arrived in the 19th century. They had a profound effect on indigenous culture. They insisted people should wear clothes to cover themselves. They eradicated many local practices and traditions. The island was made a French possession in 1853 in an attempt by Napoleon III to rival the British colonies in Australia and New Zealand. Following the example set by the British in nearby Australia, between 1854 and 1922 France sent a total of 22,000 convicted felons to penal colonies along the south-west coast of the island; this number includes regular criminals as well as political prisoners such as Parisian socialists and Kabyle nationalists. Towards the end of the penal colony era, free European settlers (including former convicts) and Asian contract workers by far out-numbered the population of forced workers. The indigenous Kanak populations declined drastically in that same period due to introduced diseases and an apartheid-like system called Code de l'Indigénat which imposed severe restrictions on their livelihood, freedom of movement and land ownership. New Caledonia has been on a United Nations list of non-self-governing territories since 1986. This list includes such places as the American Samoa, the British Falkland Islands, or the New Zealand territory of Tokelau, but noticeably it does not include places like Tibet or West Papua, which has led to its reputation as a politically biased list. Agitation by the Front de Libération Nationale Kanak Socialiste (FLNKS) for independence began in 1985. The FLNKS (led by the late Jean Marie Tjibaou, assassinated in 1989) advocated the creation of an independent state of 'Kanaky'. The troubles culminated in 1988 with a bloody hostage taking in Ouvéa. The unrest led to agreement on increased autonomy in the Matignon Accords of 1988 and the Nouméa Accord of 1998. This Accord describes the devolution process as "irreversible" and also provides for a local Caledonian citizenship, separate official symbols of Caledonian identity (such as a "national" flag), as well as mandating a referendum on the contentious issue of independence from the French Republic sometime after 2014.

Politics

Main article: Politics of New Caledonia The unique status of New Caledonia is in between that of an independent country and a regular overseas département of France. On the one hand, a territorial Congress (Congrès du territoire) and a government of the territory have been established, and a devolution of powers is organized by the 1998 Nouméa Accord. Key areas such as taxation, labor law, health and hygiene and foreign trade are already in the hands of the territorial Congress. Further competence will supposedly be given to the territorial Congress in the near future. Eventually, the French Republic should only remain competent for foreign affairs, justice, defence, public order, and treasury. A New Caledonian "citizenship" has also been introduced: only New Caledonian "citizens" have the right to vote in the local elections. This measure has been criticized, because it creates a second-class status for French citizens living in New Caledonia who do not possess New Caledonian "citizenship" (because they settled in the territory recently). New Caledonia is also allowed to engage in international cooperation with independent countries of the Pacific Ocean. Finally, the territorial Congress is allowed to pass statutes that are derogatory to French law in a certain number of areas. On the other hand, New Caledonia remains an integral part of the French Republic. Inhabitants of New Caledonia are French citizens and carry French passports. They take part in the legislative and presidential French elections. New Caledonia sends two representatives to the French National Assembly and one senator to the French Senate. The representative of the French central state in New Caledonia is the High Commissioner of the Republic (Haut-Commissaire de la République, locally known as "haussaire"), who is the head of civil services, and who seats in the government of the territory. It was decided in the Nouméa Accord that the territorial Congress will have the right to call for a referendum on independence after 2014, at a time of its choosing. The current president of the government elected by the territorial Congress is Marie-Noëlle Thémereau, from the loyalist (i.e. anti-independence) Avenir ensemble party ("Future together"), which toppled the long-time ruling RPCR (Rally for Caledonia inside the Republic) in May 2004. "Future Together" is a party of mostly Caucasian and Polynesian New Caledonians opposed to independence but tired of the hegemonic and allegedly corrupt anti-independence RPCR. Their toppling of the RPCR (that was until then seen as the only voice of New Caledonian whites) was a surprise to many, and a sign that the society of New Caledonia is undergoing changes. "Future together", as the name implies, is opposed to a racial vision of New Caledonian society, opposing Melanesians native inhabitants and European settlers, and is in favor of a multicultural New Caledonia, better reflecting the existence of large populations of Polynesians, Indonesians, Chinese, and other immigrants. Some members of "Future Together" are even in favor of independence, though not necessarily on the same basis as the Melanesian independence parties.

Demographics

Main article: Demographics of New Caledonia Though still the relative majority the indigenous Melanesian Kanak community now represents 42.5% of the population (as of 2005). This is due to a population decline during the early stages of the colonisation process. Later it was the immigration under French rule that influenced the balance in favour of other minority groups. The current population is made up of numerous ethnic groups who arrived in New Caledonia over the past 200 years: Whites (37.1%), Polynesians (Wallisians, Tahitians) (12.2%), Javanese (3.6%), Vietnamese (1.6%), and various other groups such as ni-Vanuatu, Malabaris, Maghrebins, Japanese, Chinese, Fijians (3.0%). Some of this immigration was a direct consequence of various conflicts around the world but in particular of the crumbling of the French Empire. The Kanak are known officially as Melanesians. Similarly, those whose roots are in French Polynesia are known either as Tahitians (which excludes persons originating in the other archipelagoes of French Polynesia) or simply as Polynesians (which would include both Tahitians and Wallisians, as well as many other minor groups). Whites that have lived in New Caledonia for several generations are locally known as "Caldoches", whereas newcomers who have immigrated from metropolitan France are called "Métros" or "Métropolitains". Within the official statistical category "Europeans" no distinction is made between Caledonian-born whites and French-born whites, however it is estimated that approximately two thirds identify themselves with the Caldoche community while the rest see themselves primarily as French immigrants. There is a significant contingent of people that arrive from France to work for a year or two and others that have come to retire. The Caldoche usually refer to themselves simply as "calédoniens" and may be either white (mostly French or German) or white with an admixture of Asian, Melanesian or Polynesian ancestry. Caldoche culture has many similarities with Australian and Afrikaner culture. Until very recently the Kanak population held an economically disadvantaged position in New Caledonian society, while wealthy French expatriats formed the top of the socio-economic hierarchy. The Asian and Polynesian inhabitants dominate certain segments of the local economy. There have been frequent accusations by the pro-independence movement that the French government is attempting to skew the demographic balance between the ethnic communities by clandestinely settling thousands of people from mainland France among the white Caledonians. They argue that current French policies in New Caledonia mimic those of Turkey, which has for many years been secretly chanelling in Anatolian Turks settlers to assimilate into the Turkish Cypriot population of Northern Cyprus for many years. Censuses are extremely critical to the balance of power in New Caledonia, and the organization of a new census was regularly postponed after 1996. Eventually the census was carried out in August and September 2004, amidst raging controversies over ethnic questions. Due to an intervention by French president Jacques Chirac, questions asking for the ethnicity of people were deleted from the 2004 census, officially because they were deemed to contravene the French Constitution, which states that no distinction based on ethnicity or religion should be made among French citizens. The indigenous Melanesian Kanak leaders, who are extremely sensitive to ethnic balance issues, called for New Caledonians of Kanak ethnicity not to return census forms if questions regarding ethnicity were not asked, threatening to derail the census process. Eventually, the stalemate was resolved when the local New Caledonian statistical office (a branch of the national French statistical office INSEE) agreed to ask questions regarding ethnicity. However, it is not known whether questions regarding ethnicity were asked to all residents of New Caledonia, and at any rate no data have been released, leaving the ethnic tables from the 1996 census as the only information on ethnicity currently available. According to the 2004 census, there were 230,789 inhabitants in New Caledonia as of September 2004. This means a population increase of 1.9% per year since 1996, which is less than had been anticipated. In particular, Kanak leaders were fearful of a large influx of white people from metropolitan France which would alter the ethnic balance in the territory. Census results show that immigration to New Caledonia was not as high as anticipated, with nonetheless a positive migration flow of about 1,000 people yearly to New Caledonia between 1996 and 2004 (i.e. there are 1,000 more people who migrate to New Caledonia every year than people who leave New Caledonia).

Miscellaneous

- Communications in New Caledonia
- Disputes - international: Matthew and Hunter Islands claimed by France and Vanuatu.
- Economy of New Caledonia
- Military of New Caledonia
- Music of New Caledonia
- Sister city: Gold Coast, Australia
- Transportation in New Caledonia

Publications

- J. W. Anderson, Fiji and New Caledonia, (London, 1880)
- Alfred Schreiner, La Nouvelle-Calédonie depuis sa découverte jusqu'à nos jours, (Paris, 1882)
- León Vallée Essai d'une bibliographie de la Nouvelle-Cadédonie et dépendances, (Paris, 1885)
- G. Griffith, In an Unknown Prison Land: An Account of Convicts and Colonists in New Caledonia, (London, 1901)
- J. B. Alberti, La colonisation à la Nouvelle-Calédonie, (Paris, 1909)

External links and references

- [http://www.endemia.nc Endemic flora of New-Caledonia] (French language)
- [http://www.southpacific.org/text/new_caledonia.html Finding New Caledonia]
- [http://members.ozemail.com.au/~rodeime/newcal/ "France's Best Kept Secret"] by Roderick Eime
- [http://www.janeresture.com/newcal/index.htm Jane's New Caledonia Home Page]
- [http://www.mapsouthpacific.com/new_caledonia/index.html Map of New Caledonia]
- [http://www.virtualoceania.net/newcaledonia/photos/ New Caledonia photos]
- [http://www.pacific-pictures.com/new_caledonia/index.html New Caledonia travel photos]
- [http://www.photos-nouvelle-caledonie.com/ New-Caledonia pictures] 500 high-quality pictures of New-Caledonia (French language)
- [http://www.kanaky-nouvelle-caledonie.com/ Tourism Information of New-Caledonia] (French language)
- [http://www.anytravels.com/australia/caledonia/ New Caledonia Travel Overview]
- This article incorporates edited material from the CIA World Factbook 2000
- [http://www.willgoto.com/categories.aspx?Destination=340&Langue=1 Willgoto New Caledonia] Travel guide and directory
- [http://www.vie-publique.fr/decouverte_instit/instit/instit_3_7_0_q0.htm past and current developments of France's overseas administrative divisions like New Caledonia (French language)]
- [http://www.croixdusud.info/eng/acc_eng/nc_acc_eng.php Croix du Sud] Good source of information in English and French
- [http://www.brousse-en-folie.com/broussefolie/dico/dic_acc.php Dictionnaire Franco-Calédonien de brousse-en-folie.com] A comprehensive list of French words, idioms and phrases unique to New Caledonia Category:Melanesia Category:New Caledonia Category:Special territories of the EU zh-min-nan:Sin Calédonie ja:ニューカレドニア

DNA

:For other uses, see DNA (disambiguation). DNA (disambiguation) Deoxyribonucleic acid (DNA) is a nucleic acid that contains the genetic instructions specifying the biological development of all cellular forms of life (and most viruses). DNA is a long polymer of nucleotides and encodes the sequence of the amino acid residues in proteins using the genetic code, a triplet code of nucleotides. In complex cells (eukaryotes), such as those from plants, animals, fungi and protists, most of the DNA is located in the cell nucleus. By contrast, in simpler cells called prokaryotes (the eubacteria and archaea), DNA is not separated from the cytoplasm by a nuclear envelope. The cellular organelles known as chloroplasts and mitochondria also carry DNA. DNA is often referred to as the molecule of heredity as it is responsible for the genetic propagation of most inherited traits. These traits can range from hair colour to disease susceptibility. During cell division, DNA is replicated and can be transmitted to offspring during reproduction. Lineage studies can be done based on the facts that the DNA in mitochondria (mitochondrial DNA) only comes from the mother, and the male "Y" chromosome only comes from the father. Every person's DNA, their genome, is inherited from both parents. The mother's mitochondrial DNA together with twenty-three chromosomes from each parent combine to form the genome of a fertilized egg. As a result, with certain exceptions such as red blood cells, most human cells contain 23 pairs of chromosomes, together with mitochondrial DNA inherited from the mother.

DNA Overview

red blood cell This section presents an introductory and therefore incomplete overview of DNA.
- Genes can be loosely viewed as the organism's "cookbook" or "blueprint";
- A strand of DNA contains genes, areas that regulate genes, and areas that either have no function, or a function we do not (yet) know (also see last bullet point in this section for the difference between DNA and RNA);
- DNA is organized as two complementary strands, head-to-toe, with bonds between them that can be "unzipped" like a zipper, separating the strands;
- DNA is a chain of chemical "building blocks", called "bases", of which there are four types: these can be abbreviated A, T, C, and G. Each base can only "pair up" with one single predetermined other base: A+T, T+A, C+G and G+C are the only possible combinations; that is, an "A" on one strand of double-stranded DNA will "mate" properly only with a "T" on the other, complementary strand;
- N.B.: U occasionally replaces T, notably in PBS1 phage DNA; you can thus substitute "U" for "T" throughout this section.
- Because each strand of DNA has a directionality, the sequence order does matter: A+T is not the same as T+A, just as C+G is not the same as G+C;
- For each given base, there is just one possible complementary base, so naming the bases on the conventionally chosen side of the strand is enough to describe the entire double-strand sequence;
- The genetic information contained in a strand of DNA is determined by the sequence of bases along its length;
- The cell begins DNA replication by forcibly unzipping the DNA double strand down the middle, and then recreates the "other half" of each new single strand by drowning each half in a "soup" made of the four bases. An enzyme makes a new strand by finding the correct "base" in the soup and pairing it with the original strand. In this way, the base on the old strand dictates which base will be on the new strand, and the cell ends up with an extra copy of its DNA.
- Mutations are simply chemical imperfections in this process: a base is accidentally skipped, inserted, or incorrectly copied, or the chain is trimmed, or added to; many basic mutations can be described as combinations of these accidental "operations". Mutations can also occur through chemical damage (through mutagens), light (UV damage), or through other more complicated gene swapping events.
- DNA (for DeoxyriboNucleic Acid) differs from RNA (for RiboNucleic Acid) by having the sugar 2-deoxyribose instead of ribose in its backbone (ribose contains one extra oxygen atom compared to deoxyribose -- in other words, DNA contains deoxygenated ribose, whereas RNA contains "plain" ribose.) This is the basic chemical distinction between RNA and DNA.

DNA in practice

DNA in crime

Forensic scientists can use DNA located in blood, semen, skin, saliva, or hair left at the scene of a crime to identify a possible suspect, a process called genetic fingerprinting or DNA profiling. In DNA profiling the relative lengths of sections of repetitive DNA, such as short tandem repeats and minisatellites, are compared. DNA profiling was developed in 1984 by English geneticist Alec Jeffreys, and was first used in 1986 in the Enderby murders case in Leicestershire, England. Many jurisdictions require convicts of certain types of crimes to provide a sample of DNA for inclusion in a computerized database. This has helped investigators solve old cases where the perpetrator was unknown and only a DNA sample was obtained from the scene (particularly in rape cases between strangers). This method is one of the most reliable techniques for identifying a criminal, but is not always perfect, for example if no DNA can be retrieved, or if the scene is contaminated with the DNA of several possible suspects.

DNA in computation

Despite its biological origins, DNA plays an important role in computer science, both as a motivating research problem and as a method of computation in itself, called DNA computing. As a simple example, research on string searching algorithms, which find an occurrence of a sequence of letters inside a larger sequence of letters, was motivated by DNA research, where it is used to find specific sequences of nucleotides in a large sequence. In other applications like text editors, even simple algorithms for this problem usually suffice, but DNA sequences cause these algorithms to exhibit near-worst-case behavior due to their small number of distinct characters. Databases have also been strongly motivated by DNA research, which requires special tools for storing and manipulating DNA sequences. Databases specialized for this purpose are called genomic databases, and have a number of unique technical challenges associated with the operations of approximate matching, sequence comparison, finding repeating patterns, and homology searching. In 1994, Leonard Adleman of the University of Southern California made headlines when he discovered a way of solving the directed Hamiltonian path problem, an NP-complete problem, using tools from molecular biology, in particular DNA. The new approach, dubbed DNA computing, has practical advantages over traditional computers in power use, space use, and efficiency, due to its ability to highly parallelize the computation (see parallel computing)(there is labor worth mention involved in retrieving answers computed these computational DNA techniques.). A number of other problems, including simulation of various abstract machines, the boolean satisfiability problem, and the bounded version of the Post correspondence problem, have since been analyzed using DNA computing. Due to its compactness, DNA also has an important role in cryptography, where in particular it allows unbreakable one-time pads to be efficiently constructed and used.[http://citeseer.ist.psu.edu/gehani99dnabased.html]

Overview of molecular structure

one-time pad Although sometimes called "the molecule of heredity", pieces of DNA as people typically think of them are not single molecules. Rather, they are pairs of molecules, which entwine like vines to form a double helix (see the illustration at the right). Each vine-like molecule is a strand of DNA: a chemically linked chain of nucleotides, each of which consists of a sugar, a phosphate and one of five kinds of nucleobases ("bases"). Because DNA strands are composed of these nucleotide subunits, they are polymers. The diversity of the bases means that there are five kinds of nucleotides, which are commonly referred to by the identity of their bases. These are adenine (A), thymine (T), uracil (U), cytosine (C), and guanine (G). U is rarely found in DNA except as a result of chemical degradation of C, but in some viruses, notably PBS1 phage DNA, U completely replaces the usual T in its DNA. Similarly, RNA usually contains U in place of T, but in certain RNAs such as transfer RNA, T is always found in some positions. Thus, the only true difference between DNA and RNA is the sugar, 2-deoxyribose in DNA and ribose in RNA. In a DNA double helix, two polynucleotide strands can associate through the hydrophobic effect and pi stacking. Specificity of which strands stay associated is determined by complementary pairing. Each base forms hydrogen bonds readily to only one other -- A to T and C to G -- so that the identity of the base on one strand dictates the strength of the association; the more complementary bases exist, the stronger and longer-lasting the association. The cell's machinery is capable of melting or disassociating a DNA double helix, and using each DNA strand as a template for synthesizing a new strand which is nearly identical to the previous strand. Errors that occur in the synthesis are known as mutations. The process known as PCR (polymerase chain reaction) mimics this process in vitro in a nonliving system. Because pairing causes the nucleotide bases to face the helical axis, the sugar and phosphate groups of the nucleotides run along the outside; the two chains they form are sometimes called the "backbones" of the helix. In fact, it is chemical bonds between the phosphates and the sugars that link one nucleotide to the next in the DNA strand.

The role of the sequence

Within a gene, the sequence of nucleotides along a DNA strand defines a messenger RNA sequence which then defines a protein, that an organism is liable to manufacture or "express" at one or several points in its life using the information of the sequence. The relationship between the nucleotide sequence and the amino-acid sequence of the protein is determined by simple cellular rules of translation, known collectively as the genetic code. The genetic code is made up of three-letter 'words' (termed a codon) formed from a sequence of three nucleotides (e.g. ACT, CAG, TTT). These codons can then be translated with messenger RNA and then transfer RNA, with a codon corresponding to a particular amino acid. There are 64 possible codons (4 bases in 3 places

4^3

) that encode 20 amino acids. Most amino acids, therefore, have more than one possible codon. There are also three 'stop' or 'nonsense' codons signifying the end of the coding region, namely the UAA, UGA and UAG codons. In many species, only a small fraction of the total sequence of the genome appears to encode protein. For example, only about 1.5% of the human genome consists of protein-coding exons. The function of the rest is a matter of speculation. It is known that certain nucleotide sequences specify affinity for DNA binding proteins, which play a wide variety of vital roles, in particular through control of replication and transcription. These sequences are frequently called regulatory sequences, and researchers assume that so far they have identified only a tiny fraction of the total that exist. "Junk DNA" represents sequences that do not yet appear to contain genes or to have a function. The reasons for the presence of so much non-coding DNA in eukaryotic genomes and the extraordinary differences in genome size ("C-value") among species represent a long-standing puzzle in DNA research known as the "C-value enigma". Some DNA sequences play structural roles in chromosomes. Telomers and centromeres typically contain few (if any) protein-coding genes, but are important for the function and stability of chromosomes. Some genes code for "RNA genes" (see tRNA and rRNA). Some RNA genes code for transcripts that function as regulatory RNAs (see siRNA) that influence the function of other RNA molecules. The intron-exon structure of some genes (such as immunoglobin and protocadeherin genes) is important for allowing alternative splicing of pre-mRNA which allows several different proteins to be made from the same gene. Some non-coding DNA represents pseudogenes that can be used as raw material for the creation of new genes with new functions. Some non-coding DNA provided hot-spots for duplication of short DNA regions; such sequence duplication has been the major form of genetic change in the human lineage (see evidence from the Chimpanzee Genome Project). Exons interspersed with introns allows for "exon shuffling" and the creation of modified genes that might have new adaptive functions. Large amounts of non-coding DNA is probably adaptive in that it provides chromosomal regions where recombination between homologous portions of chromosomes can take place without disrupting the function of genes. Some biologists such as Stuart Kauffman have speculated that there must be mechanisms by which the rate of evolution of a species can be increased or decreased. Non-coding DNA provides mechanisms for gene creation, modification and recombination it is probably important for control of the rate of human evolution. Sequence also determines a DNA segment's susceptibility to cleavage by restriction enzymes, the quintessential tools of genetic engineering. T